Since their discovery in 1977 hydrothermal vent communities have provided a wealth of surprises about life in the deep-sea and in extreme environments. In contrast to most other marine ecosystems which largely depend on photosynthesis, these faunal communities are characterized by their dependence on geochemical energy sources (largely hydrogen sulfide and methane). These faunas pose numerous questions about how and when they adapted to these environments, and how they survive under such extreme conditions. Soon after the discovery of deep-sea vent faunas, ecosystems with a similar suite of taxa were found at hydrocarbon seeps in the deep-sea (cold-seeps), at sunken whale carcasses (whale-falls), and sunken driftwood (wood-falls). Such environments are now known as ‘chemosynthetic ecosystems’.